Abstract of Stearley, R.F., and G. R. Smith. 1993. Phylogeny of the Pacific trouts and salmons (Oncorhynchus) and genera of the family Salmonidae. Trans. Amer. Fish. Soc. 122(1):1-33.

The phylogeny of the salmonidae shows lake Ohrid forms to be primitive, near Brachymystax. Chars are more advanced; the most advanced genera are Oncorhynchus, the pacific trout and salmon, and Salmo, the atlantic trout and salmon.