Abstract of Smith, G.R., and T. N. Todd. 1992. Morphological cladistic study of coregonine fishes. In T. N. Todd and M. Luczynski (Eds.) Proc. Internat. Symp. Biol. Manag. Coregonid Fishes. Univ. Agr. Tech., Olstzyn - Kortowo, Poland. 39:479-490.

Prosopium is the most primitive group of whitefishes. Among the species of Coregonus, whitefish and ciscoes are different clades with different species groups. The inconu, "Stenodus" is a cisco.