Class Elasmobranchomorpha--sharks, rays, skates, chimaeras and Iniopterygians probably sister group to Osteichthys (Young 86, Money 84, Schaeffer 75), based on long palatoquadrate, expanded posteriorly teeth develop from dental lamina subclass Chondrichys--sharks, skates, and rays probably a natural group subclass Holocephali--chimaeras, ratfish uncertain affinities, Chondrichthys or Ptyctodonts Chondrichthyan denticles (and fish scales) in Upper Silurian good Chondrichthys first appear in late lower Devonian e.g., Cladodus M Dev --Perm; Cladoselachii Upper Devonian to Pennsylvanian; Selachians with Ctenochanthus (Penn), Hybodus (Perm- Cret), Symmoriids (plesiomorphic group); Characters: no swimbladder no internal or external bony skeleton (calc. cartilage) heterocercal caudal? paired fins P1 P2 D 1 or 2 (w spines after Paleozoic), A later, fins with basals and radials oviparous or viviparous claspers in males placoid teetn drrived from scales, dentine and enamel-like two ancient branches subterbranchialia (iniopterygia, chondrenchelyidae, helodontidae, menaspidae, cochliodontidae, chimaridae, ) Elasmobranchii 2 branches: Euselachii-- (etc, Xenacanthida, cladoselachida, symmoriidae, orodontids, petalodontids +_ orodontida 7 major Paleozoic groups cladoselachida USil Dev, Dspines, e.g. Cladoselache symmorids--c symmetrical Eugeniodontids--symphyseal whorl Squatinactida--raylike Petalodontids--shark like Xenacanthida--Dev Triass, 2 A fins Ctenacanthida>>Hybodontids (&Neoselachii)? mostly marine history