Abstract of Dowling, T.E., W.R. Hoeh, G.R. Smith, and W. M. Brown. 1992. Evolutionary relationships of shiners in the genus Luxilus, (Cyprinidae) as determined by analysis of mitochondrial DNA. Copeia 1992:306-322.

  • ((((cornutus, albeolus)(isolepis, chrysocephalus)) (coccogenis, zonistius) (cardinalis-pilsbryi, zonatus)) cerasinus)